After a century of hypotheses without conclusive evidence, well-preserved fossils of feathered dinosaurs were discovered during the 1990s, and more continue to be found. The fossils were preserved in a Lagerstätte a sedimentary deposit exhibiting remarkable richness and completeness in its fossils in Liaoning, China. The area had repeatedly been smothered in volcanic ash produced by eruptions in Inner Mongolia 124 million years ago, during the Early Cretaceous Period. The fine-grained ash preserved the living organisms that it buried in fine detail. The area was teeming with life, with millions of leaves, angiosperms (the oldest known), insects, fish, frogs, salamanders, mammals, turtles, and lizards discovered to date.
The most important discoveries at Liaoning have been a host of feathered dinosaur fossils, with a steady stream of new finds filling in the picture of the dinosaurbird connection and adding more to theories of the evolutionary development of feathers and flight. Norell et al. (2007) reported quill knobs from an ulna of Velociraptor mongoliensis, and these are strongly correlated with large and well-developed secondary feathers.
A nesting Citipati osmolskae specimen, at the AMNH.
Behavioural evidence, in the form of an oviraptorosaur on its nest, showed another link with birds. Its forearms were folded, like those of a bird.[6] Although no feathers were preserved, it is likely that these would have been present to insulate eggs and juveniles.